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Chris Lloyd Mills

Senior Lecturer

School of Science & Technology

Staff Group(s)


Module Leaderships in Practical Techniques in:

  • Biology (level 1)
  • Dugs and Tissues (level 2)
  • Applied Molecular Physiology (level 2)
  • Cellular Physiology and Toxicology (level 3)

Teaching contributions to other modules are:

  • Essential of Living Systems (level 1)
  • Human Systems (level 1)
  • Introduction to Pharmacology (level 1)
  • Integrative Human Physiology (level 2)
  • Current Topics in Pharmacology (level 3)
  • Current Topics in Physiology (level 3)
  • Pharmacology (Masters level)

Research areas

Dr Lloyd Mills main areas of research interest are:

  • Crustacean Ecophysiology
    • Physiological environmental adaptations
    • Invasive species
    • aquatic pollution
  • Cellular Physiology
    • Mechanisms of heavy metal toxicity
    • Calcium signalling

Opportunities to carry out postgraduate research towards an MPhil / PhD exist in all the areas identified above and in associated interdisciplinary areas. Further information may be obtained from the NTU Graduate School.

External activity

Sponsors and collaborators

  • Royal Society
  • Dr Jamie Dick (Queens University, Belfast)
  • Dr Alison Dunn (Leeds University)
  • Dr Dirk Plavoet (University of Amsterdam, the Netherlands)
  • Dr Paul Wood (Loughborough University)


Selected publications

  • Parasite infection manipulates sodium regulation in the freshwater amphipod Gammarus pulex (L.). Brooks SJ and Lloyd Mills C, Comparative Biochemistry and Physiology A: Molecular and Integrative Physiology, 2011, 159, 247-252, DOI: 10.1016/j.cbpa.2011.03.011
  • Osmoregulation in the hypogean freshwater amphipod, Gammarus pulex (L.). Brooks SJ and Lloyd Mills C, Journal of Crustacean Biology, 2011, 31, 332-338, DOI: 10.1651/10-3336.1
  • Cation regulation and alteration of water permeability in the amphipod Dikerogammarus villosus: an indicator of invasion potential. Brooks SJ, Platvoet D and Lloyd Mills C, Fundamental and Applied Limnology, 2008, 172, 183-189. DOI: 10.1127/1863-9135/2008/0172-0183
  • Reduced tubulin tyrosination as an early marker of mercury toxicity in differentiating N2a cells. Lawton M, Iqbal M, Kontovraki M, Lloyd Mills C and Hargreaves AJ, Toxicology in vitro, 2007, 21 (7), 1258-61
  • Gill Na+, K+-ATPase in a series of hyper-regulating gammarid amphipods. Enzyme characterisation and the effects of salinity acclimation. Brooks SJ and Lloyd Mills C, Comp. Biochem. Physiol. 2006, 144A, 24-32
  • Development of a new low cost high sensitivity system for behavioural ecotoxicity testing. Lloyd Mills C, Shukla DH and Compton GJ, Aquatic Toxicology, 2006, 77, 197-201

See all of Chris Lloyd Mills's publications...